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GSH is regulated by GSH synthase, GSH peroxidase and GSH reductase (GR).9 GR, which is one of the key enzymes in the redox system of the human body, is ubiquitous in the liver, kidney, heart, erythrocytes and other tissue cells, and it primarily functions to convert oxidised GSH (GSSG) into reduced GSH, scavenge oxygen free radicals and protect cells from oxidative damage

229 Conversely, SIRT1 facilitates transcriptional activation of the BECN1 gene by deacetylating histone H4 at lysine 16, thereby establishing a chromatin configuration permissive for autophagy induction under nutrient-deprived conditions